Non-marine invertebrate trace fossils from the Tertiary Calatayud-Teruel basin, NE Spain

Relatively diverse trace fossils made by insects, other arthropods and oligochaete worms occur in the Miocene lacustrine marginal deposits of Calatayud-Teruel basin (NE Spain). They include ichnospecies Celliforma isp., Celliforma? isp. A B, aff. habari, Rosellichnus cf. arabicus, Spongeliomorpha Labyrintichnus terrerensis n. igen. et Taenidium barreti, Beaconites filiformis Polykladichnus aragonensis Their taxonomic ethologic interpretations cross-cutting relationships permit to envisage new lines evidence for reconstructing transitions from dry-ground terrestrial moist-ground subaqueous environments, related episodic floodings ponds. These environmental (related external controls) are characterized benthic community replacements, evinced vertical successions Termitichnus, Scoyenia Mermia-like ichnofacies.